This little oversight can be exploited: Say mario wants to collect a star to finish a level. He needs to position himself somewhat close to the star, but only his position after modulo 65536 is used for collision detection. We can use another exploit to make Mario gain massive velocity and the physics engine will allow him to clip through walls with it. However, with great velocity comes the price of leaving the map (going out of bounds). Therefor, we use the module operation to still force a collision detection with the star.
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Linda Cardellini in "DTF St. Louis."